Publications

Below is a list of my current publications and their themes. You can also see my Google Scholar, ORCiD or Web of Science profiles.

Publication themes key

Management of big data & databases
Statistical analysis
Computational methods creation

Morphology & morphometrics
Palaeontological specimens & field work
Palaeoecology & fluid dynamics
Open science principles

Scientific publications

	Cooper, N. & BES Data and Code Hackathon Group. 2026. Data‐and code‐archiving in the British Ecological Society journals: Present status and recommendations for future improvements. Methods in Ecology and Evolution 17	online access
	Lynch, S., Gueriau, P., Drage, H.B., Dutheil, D.B., Charbonnier, S., Corthésy, N., Luque, J., Martin Olmos, C. & Daley, A.C. 2026. Moulting and development in a freshwater prawn from the Cretaceous of Morocco. PeerJ 14	online access
	Daley, A.C., Gueriau, P., Lustri, L., Drage, H.B., Lynch, S., Monceret, E. & Monceret, S. 2026. Euarthropods from the Early Ordovician Cabrières Biota, including a new aglaspidid and freshly moulted trilobite material. Lethaia 59	online access
	Saleh, F., et al. 2026. Preservation modes of exceptionally preserved fossils from the Early Ordovician Cabrières Biota (France). Lethaia 59	online access
	Marty, D., Cavin, L., Daley, A.C., Drage, H.B., Hautmann, M., Joyce, W. & Klug, C. 2026. Editorial – Swiss Journal of Palaeontology migrates to Pensoft Publishers. Swiss Journal of Palaeontology 145	online access
	Leone, M., Rech de Laval, V., Drage, H.B., Waterhouse, R.M. & Robinson-Rechavi, M. 2026. TaxonMatch: taxonomic integration and tree construction from heterogeneous biological databases. bioRxiv preprint	online access
	Kim, K., Lynch, S., Drage, H.B., Antcliffe, J.B., Chipman, A.D., Daley, A.C. & Robinson-Rechavi, M. Morphological characterization of moulting in the Atlantic horseshoe crab Limulus polyphemus and evolutionary insights into ecdysis patterns in Arthropoda. 2026. bioRxiv preprint	online access
	Drage, H.B. & Pates, S. 2025. Hierarchical importance of factors impacting trilobite cephalic disparity. bioRxiv preprint	online access
	Jones L.A., Dean C.D., Allen B.J., Drage H.B., Gearty, W., Flannery-Sutherland J.T., Chiarenza A.A., Dillon E.M., Farina B.M. & Godoy P.L. Ten simple rules to follow when cleaning occurence data in palaeobiology. Palaeontology 68	online access
	Drage, H.B., Keating, J.N., Nielsen, M.L., Saleh, F. & Wong Hearing, T.W. 2024. Open Palaeontology: a new model of diamond open access journal for palaeontology. Open Palaeontology 1:1	online access
	Drage, H.B. & Pates, S. 2024. Distinct causes underlie double-peaked trilobite morphological disparity in cephalic shape. Communications Biology 7:1490	online access
	Drage, H.B., Pates, S. & Minter, N.J. 2024. Experimental protocol for validation of computational fluid dynamics palaeontological simulations. EcoEvoRxiv preprint	online access
	Pates, S. & Drage, H.B. 2024. Hydrodynamic function of genal prolongations in trinucleimorph trilobites revealed by computational fluid dynamics. BioRxiv preprint	online access
	Campli, G., Volovych, O., Kim, K., Veldsman, W.P., Drage, H.B., Sheizaf, I., Lynch, S., Chipman, A.D., Daley, A.C., Robinson-Rechavi, M. & Waterhouse, R.M. 2024. The moulting arthropod: a complete genetic toolkit review. Biological Reviews 99:6	online access
	Saleh, F., et al. 2024. Reply to: The Cabrières Biota is not a Konservat-Lagerstätte. Nature Ecology & Evolution 8	online PDF
	Lynch, S., Żylińska, A., Daley, A.C. & Drage, H.B. 2024. Dual role of enrolment for moulting and protection in a Cambrian trilobite from Poland. Lethaia 57:1	online access
	Saleh, F., et al. 2024. The Cabrières Biota (France) provides insights into Ordovician polar ecosystems. Nature Ecology & Evolution 8	online access
	Drage, H.B. 2024. Trilobite moulting behaviour variability had little association with morphometry. Palaeontologia Electronica 27(1):a9	online access PDF
	Drage, H.B. & Wong Hearing, T.W. [joint first-authors] 2023. Diamond Open Access with Preregistration: a new publishing model for palaeontology. EarthArXiv preprint	online access
	Laibl, L., Gueriau, P., Saleh, F., Pérez-Peris, F., Lustri, L., Drage, H.B., Bath Enright, O.G., Potin, G. & Daley, A.C. 2023. Immature stages of a Lower Ordovician marrellid from Morocco suggest size-induced ontogenetic niche differentiation in early euarthropods. Frontiers in Ecology & Evolution 11	online access
	Chipman, A. & Drage, H.B. [joint first-authors] 2023. Trilobites in rock enrol: Commentary on Esteve and Hughes (2023). Proceedings of the Royal Society B 290	online access
	Laibl, L., Drage, H.B., Pérez-Peris, F., Schöder, S., Saleh, F. & Daley, A.C. 2023. Babies from the Fezouata: adaptations to high latitudes in the early developmental trilobite stages. Geobios 81	online access
	Drage, H.B., Legg, D.A. & Daley, A.C. 2023. Novel marrellomorph moulting behaviour preserved in the Lower Ordovician Fezouata Shale, Morocco. Frontiers in Ecology & Evolution 11	online access
	Drage, H.B., Holmes, J.D., García-Bellido, D.C. & Paterson, J.R. 2023. Associations between trilobite intraspecific moulting variability and body proportions: Estaingia bilobata from the Cambrian Emu Bay Shale, Australia. Palaeontology 66	online access
	Feldsman, W.P., Campli, G., Dind, S., Rach de la Vale, V., Drage, H.B., Chipman, A.D., Waterhouse, R.M., & Robinson-Rechavi, M. 2022. Taxonbridge: an algorithm for the creation and analysis of custom taxonomies. BioRxiv preprint	online access
	Drage, H.B., Vandenbroucke, T., Van Roy, P. & Daley, A.C. 2019. Sequence of post-moult exoskeleton hardening preserved in a trilobite mass moult assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco. Acta Palaeontologica Polonica 64:2	PDF
	Drage, H.B. 2019. Quantifying intra- and interspecific variability in trilobite moulting behaviour across the Palaeozoic. Palaeontologia Electronica 22:2	PDF
	Yang, J., Ortega-Hernández, J., Drage, H.B., Du, K. & Zhang, X. 2019. Ecdysis in a stem-group euarthropod from the early Cambrian of China. Scientific Reports 9:1	online access
	Daley, A.C., Antcliffe, J.B., Drage, H.B. & Pates, S. 2018. Early fossil record of Euarthropoda and the Cambrian Explosion. PNAS 115:21	online access
	Drage, H.B., Laibl, L. & Budil, P. 2019. Post-embryonic development of Dalmanitina, and the evolution of facial suture fusion in Phacopina. Paleobiology 44:4.	online PDF
	Drage, H.B., Holmes, J.D., Garcia-Bellido, D.C. & Daley, A.C. 2018. Exceptionally variable trilobite moulting behaviour preserved in the Emu Bay Shale, South Australia. Lethaia 51:4	online access
	Drage, H.B. & Daley, A.C. 2016. Recognising moulting behaviour in trilobites by examining morphology, development, and preservation: Comment on Błażejowski et al. 2015. BioEssays 38:10	online access
	Daley, A.C. & Drage, H.B. [joint first-authors] 2016. The fossil record of ecdysis, and trends in the moulting behaviour of trilobites. Arthropod Structure & Development 45:2	online PDF

Data publications, datasets, outreach publications

	Jones L.A., Dean C.D., Allen B.J., Drage H.B., Gearty, W., Flannery-Sutherland J.T., Chiarenza A.A., Dillon E.M., Farina B.M. & Godoy P.L. Ten simple rules to follow when cleaning occurence data in palaeobiology: vignette.	online access
	Rech de Laval, V., Leone, M., Drage, H.B., Lynch, S., Campli, G., Dind, S., Kim, K., Volovych, O., Sheizaf, I., Chipman, A. Daley, A.C., Waterhouse, R. & Robinson-Rechavi, M. MoultDB: arthropod moulting database and website UI. https://moulting.org.	moultdb database moulting website
	Gearty, W., Jones, L.A., Dillon, E., Godoy, P., Drage, H.B., Dean, C. & Farina, B. 2024. CRAN Task View: Paleontology. R package summary for palaeontological analyses. https://cran.r-project.org/view=Paleontology	CRAN task view
	Drage, H.B. 2024. Euarthropod diversity in Pokémon: searching for the ancestral type. Journal of Geek Studies 8:1.	online access
	Pates, S. & Drage, H.B. 2024. Trilobite genal spine 3D models and Ansys (computational fluid dynamics analyses) files.	online access
	Drage, H.B. & Pates, S. 2024. Trilobite cephalon morphometric data, 2D models to produce morphometric data, and R code.	online access
	Lynch, S., Żylińska, A., Daley, A.C. & Drage, H.B. 2024. Strenuella polonica moulting and enrolment data.	online access
	Drage, H.B. & Wong Hearing, T.W. [joint first-authors] 2023. A new model for publishing. The Palaeontological Association Newsletter 114.	online access
	Drage, H.B., Holmes, J.D., García-Bellido, D.C. & Paterson, J.R. 2023. Estaingia bilobata morphometry and moulting behaviour data and R code.	online access
	Drage, H.B. 2023. Trilobite morphometry and moulting behaviour data and R code.	online access
	Feldsman, W.P., Campli, G., Dind, S., Rach de la Vale, V., Drage, H.B., Chipman, A.D., Waterhouse, R.M., & Robinson-Rechavi, M. 2022. Taxonbridge: an algorithm for the creation and analysis of custom taxonomies.	R package
	Drage, H.B. 2019. Trilobite moulting behaviour variability data and R code.	online access

I have also published 10 newsletter issues for The Palaeontological Association as the Council Newsletter Editor, which are available on the Association website.

I host my datasets and code freely open access on the Open Science Framework or Zenodo. You can use these, or get in touch if you want to discuss anything.

From 2019 to 2021 I had a career in science education publishing at Oxford University Press. In these roles I worked on many educational publications, including the following titles. Get in touch if you’re interested in what it’s like to move from academia to publishing, or in discussing science education.

• Oxford Smart Activate KS3

• Oxford University Press Oxford Revise: A Level Physics, Chemistry, and Biology

• Oxford University Press Oxford Revise: GCSE Physics, Chemistry, and Biology

• Oxford University Press Maths Skills for AS and A Level Psychology

• Oxford University Press Maths Skills for GCSE Computer Science.